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In this situation, multiple histidine residues of the octarepeats are involved in the binding reaction

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Arch Biochem Biophys 595:6163 Kannan R, Chakrabarti R, Tang D, Kim KJ, Kaplowitz N (2000) GSH transport in human cerebrovascular endothelial cells and human astrocytes: evidence for luminal localization of Na+-dependent GSH transport in HCEC